Upcoming: Dangers of limited winegrape diversity in the Anthropocene – Interactive figure of R x S studies


Zeng, Z. A. & E. M. Wolkovich. 2024. Weak evidence of provenance effects in spring phenology across Europe and North America. New Phytologist (242): 1957-1964. (pdf // supp // data)
Jones, F. A. M., Bogdanoff, C. & E. M. Wolkovich. 2024. The role of genotypic and climatic variation at the range edge: a case study in winegrapes. American Journal of Botany (111): 1. (pdf // supp // data)
Buonaiuto, D. M., Davies, T. J., Collins, S. C. & E. M. Wolkovich. 2024. Ecological drivers for flower-leaf sequences: aridity and proxies for pollinator attraction select for flowering-first in the American plums. New Phytologist (242): 2312-2321. (pdf // supp  // data)


Kharouba, H. M. & E. M. Wolkovich. Lack of evidence for the match-mismatch hypothesis across terrestrial trophic interactions. Accepted. Ecology Letters. (pdf // supp // data)
Rivest, S. A., Wolkovich, E. M. & H. M. Kharouba. 2023. Flowering phenology influences butterfly nectar foraging on non-native plants in an oak savanna. Ecology (104): e4004. (pdf//data)
Chamberlan, C. J. & E. M. Wolkovich. 2023. Variation across space, species and methods in models of spring phenology. Climate Change Ecology (5): 100071. (pdf)
Buonaiuto, D. M. , Donahue, M. J. & E. M. Wolkovich. 2023. Experimental designs for testing the interactive effects of temperature and light in ecology and the problem of periodicity. Functional Ecology (37): 1747-1756 (pdf)
Buonaiuto, D. M. & E. M. Wolkovich. 2023. Contrasting responses to climate variability generate seasonal priority effects between native and invasive forest herbs. Journal of Ecology (111): 1711-1721. (pdf//data)
Jones, F. A. M., Bogdanoff, C., & E. M. Wolkovich. 2023. The role of genotypic and climatic variation at the range edge: A case study in winegrapes. American Journal of Botany (111): 1-12. (pdf)


Wolkovich, E. M., Chamberlain*, C.J., Buonaiuto*, D. M., Ettinger*, A.K. & I. Morales-Castilla.* 2022. How interactive effects of temperature and photoperiod shape plant phenology responses to warming. New Phytologist: 235: 1719-1728. (pdf // supp)
Ettinger*, A.K, Chamberlain*, C.J., & E. M. Wolkovich. 2022. The increasing relevance of phenology to conservation. Nature Climate Change (12): 305-309. (Invited, not peer-reviewed) (pdf)
Wolkovich, E. M., Chamberlain, C.J., Buonaiuto, D. M., Ettinger, A.K. & I. Morales-Castilla. 2022. Integrating experiments to predict interactive cue effects on spring phenology with warming. New Phytologist: 235: 1719-1728. (pdf//data)


Wolkovich, E. M., Auerbach, J. A., Chamberlain, C.J., Buonaiuto, D. M., Ettinger, A.K., Morales- Castilla, I. & A. Gelman. 2021. A simple explanation for declining temperature sensitivity with warming. Global Change Biology: 10.1111/gcb.15746. (pdf // supp // data // blog // theme song: Birdhouse in Your Soul by They Might Be Giants)
Wolkovich, E. M. & Donahue, M. 2021. How phenological tracking shapes species and communities in non-stationary environments. Biological Reviews: 10.1111/brv.12781. (pdf // supp)
Eyster, H. & Wolkovich, E. M. Comparisons in the native and introduced ranges reveal little evidence of climatic adaptation in germination traits. Accepted at Climate Change Ecology. (pdf // supp // data)
Reimer, J. R., Arroyo-Esquivel, J. Jiang, J., Scharf, E. M. Wolkovich, Zhu, K. & C. Boettiger. 2021. Noise can create or erase long transient dynamics. Theoretical Ecology: (pdf // supp // data)
Buonaiuto, D. M. & E. M. Wolkovich. 2021. Differences between flower and leaf phenological responses to environmental variation drive shifts in spring phenological sequences of temperate woody plants. Journal of Ecology: (109): 2922–2933. (pdf // supp // data)
Chamberlain, C.J & E. M. Wolkovich. 2021. Late spring freezes coupled with warming winters alter temperate tree phenology and growth. New Phytologist: (231) 987–995. (pdf // supp // data // theme song: Ice Wine by Lia Ices)
I. Morales-Castilla & E.M. Wolkovich. 2021. Diversity may save wines from climate change. The Science Breaker. (non-peer-reviewed) (article)


Ettinger, A.K., Buonaiuto, D. M., Chamberlain, C.J., Morales-Castilla & E. M. Wolkovich. Spatial and temporal shifts in photoperiod with climate change. New Phytologist: 230 (2), 462-474. (pdf // supp // data)
Ettinger, A.K., Chamberlain, C.J., Buonaiuto, D. M., Morales-Castilla, I., Flynn, D.F.B., Savas, T., Samaha, J. & E. M. Wolkovich. 2020. Winter temperatures predominate in spring phenological responses to warming. Nature Climate Change: 10 (12), 1137-1142. (pdf // supp // data // theme song: Out Of The Woods by Taylor Swift)
Chamberlain, C.J., Cook, B.I., Morales-Castilla, I. & E. M. Wolkovich. 2020. Climate change reshapes the drivers of false spring risk across European trees. New Phytologist: 229 (1), 323-334. (pdf // supp // theme song: Winter Winds by Mumford & Sons)
Merrill, N.K., I., García de Cortazar-Atuari, I., T., Parker, A., Walker, M. A. & E. M. Wolkovich. 2020. Exploring grapevine phenology and high temperatures response under controlled conditions. Frontiers in Environmental Science. (pdf // lab-data // field-data)
Buonaiuto, D. M., Morales-Castilla, I., E. M. Wolkovich. 2020. Reconciling competing hypotheses regarding flower–leaf sequences in temperate forests for fundamental and global change biology. New Phytologist. (pdf // supp // data)
Kharouba, H.M., Wolkovich, E.M. Disconnects between ecological theory and data in phenological mismatch research. Nat. Clim. Chang. 10, 406–415 (2020). (pdf // supp //data)
Morales-Castilla, I., García de Cortazar-Atuari, I., Cook, B. I., Lacombe, T., Parker, A. van Leeuwen, C. & E. M. Wolkovich. 2020. Cultivar diversity buffers winegrowing regions from climate change losses. PNAS: 117 (6), 2864-2869. (pdf // supp // data)


Ettinger,A. K., Chuine, I., Cook, B. I., Dukes, J. S., Ellison, A. M., Johnston, M. R., Panetta, A. M. Rollinson, C. R. 10 Y. Vitasse, Y. & E. M. Wolkovich. How do climate change experiments alter plot-scale climate? Ecology Letters, 2019 (22) 748–763 (pdf)
Chamberlain, C. J., Cook, B. I., Garcia de Cortazar Atauri, I. & E. M. Wolkovich. 2019. Rethinking False Spring Risk. Global Change Biology. (pdf // theme song: Shots by Imagine Dragons)
E.M. Wolkovich & I. Morales-Castilla*. 2019. Climate change: Why varietal diversity is critical to wine- growing’s warmer future. Wine & Viticulture Journal 34(1). (non-peer-reviewed) (pdf)
Joly, S, D. F. B. Flynn & E. M. Wolkovich. On the importance of accounting for intraspecific genetic correlations in multi-species studies. Methods in Ecology & Evolution. (bioRxiv // pdf)
Davies, T. J., Regetz, J., Wolkovich, E. M. & McGill, B. J. Phylogenetically weighted regression: A method for modelling non‐stationarity on evolutionary trees. Global Ecology & Biogeography. 2019: 28(2) 275-285. (pdf)


Wolkovich, E. M., I. García de Cortazar-Atauri, Morales-Castilla, I., K. A. Nicholas & T. Lacombe. 2018. From Pinot to Xinomavro in the world’s future winegrowing regions. Nature Climate Change 8:29-37. (media // pdf)
Flynn, D. F. B. & E.M. Wolkovich. 2018. Temperature and photoperiod drive spring phenology across all species in a temperate forest community. New Phytologist 219(4): 1353-1362. (pdf // theme song: Shots by Imaine Dragons)
Ettinger, A., S. Gee & E. M. Wolkovich. 2018. Phenological sequences: how early-season events define those that follow. American Journal of Botany Oct; 105(10):1771-1780. (pdf)
Kharouba, H. M., Ehrlen, J., Gelman, A., Bolmgren, K., Allen, J. M., Travers, S. & E. M. Wolkovich. Global shifts in the phenological synchrony of species interactions over recent decades. PNAS: Apr 2018, DOI:10.1073/pnas.1714511115 (pdf)


Wolkovich, E. M., D. O. Burge, M. A. Walker & K. A. Nicholas. 2017. Phenological diversity provides opportunities for climate change adaptation in winegrapes. Journal of Ecology 105:905-912. (data // pdf)


Cook, B. I. & E. M. Wolkovich. 2016. Climate change decouples drought from early winegrape harvests in France. Nature Climate Change 6:715-719. (pdf//media)
Wolkovich, E. M. Reticulated channels in soil food webs. Soil Biology and Biochemistry: 102: 18-21. (pdf)


Seabloom, E. W. & 66 co-authors (Wolkovich, E. M. co-author). 2015. Plant species’ origin predicts dominance and response to nutrient enrichment and herbivores in global grasslands. Nature Communications 6: 7710, doi:10.1038/ncomms8710. (article // pdf)


Wolkovich, E. M., Cook, B. I., McLauchlan, K. K. & T. J. Davies. 2014. Temporal ecology in the Anthropocene. Ecology Letters 17(11): 1365–1379. (pdf // theme song: Lakehouse by Of Monsters and Men)
Wolkovich, E. M., Allesina, S., Cottingham, K. L., Moore, J. C. & C. de Mazancourt. 2014. Linking the green and brown worlds: The prevalence and effect of multi-channel feeding in food webs. Ecology, 95(12), 2014, pp. 3376–3386. (pdf)
Francis, T. B., Wolkovich, E. M., Scheuerell, M. D., Katz, S. L., Elizabeth E. E., & S. E. Hampton. In press. 2014. Shifting regimes and changing interactions in the Lake Washington, U.S.A., plankton community from 1962-1992. PLoS ONE 9(10): e110363. doi:10.1371/journal.pone.0110363 (pdf)
Wolkovich, E. M. & A. K. Ettinger. 2014. Back to the future for plant phenology research (Commentary). New Phytologist 203: 1021–1023. (pdf)
Wolkovich, E. M. , B. I. Cook & T. J. Davies. 2014. Progress towards an interdisciplinary science of plant phenology: Building predictions across space, time and species diversity. New Phytologist 201: 1156–1162. (pdf // data)
Wolkovich, E. M. & E. E. Cleland. 2014. Phenological niches and the future of invaded ecosystems with climate change. AoB Plants doi:10.1093/aobpla/plu013. (pdf)


Wolkovich, E. M., Davies, T. J., Schaefer, H., Cleland, E. E., Cook, B. I., Travers, S. E. , Willis, C. G. & C. C. Davis. 2013. Temperature-dependent shifts in phenology contribute to the success of exotic species with climate change. American Journal of Botany 100(7): 1407-1421. (pdf // data)
Pau, S., Wolkovich, E. M., Cook, B. I., Nytch, C., Regetz, J., Zimmerman, J. K. & S. J. Wright. 2013. Clouds and temperature drive dynamic changes in tropical flower production. Nature Climate Change 3: 838-842. (article // HGB’s Jump in the line)
Davies, T.J., E. M. Wolkovich & 16 other co-authors. 2013. Phylogenetic conservatism in plant phenology. Journal of Ecology 101: 1520–1530. (pdf)


Wolkovich, E.M. & 18 co-authors. 2012. Warming experiments underpredict plant phenological responses to climate change. Nature 485 (7399): 494-497. (pdf // supp, data from NECTAR & STONE,  Article and Forum)
Wolkovich, E. M., Regetz, J. & M. I. O’Connor. 2012 Advances in global change research require open science by individual researchers. Global Change Biology 18(7): 2102-2110. (pdf)
Cook, B.I., Wolkovich, E.M. & C. Parmesan. 2012 Divergent responses to spring and winter warming drive community level flowering trends. Proceedings of the National Academy of Sciences 109(3): 9000-9005. (article)
Cleland, E. E., J. M. Allen, T. M. Crimmins, J. A. Dunne, S. Pau, S. E. Travers, E. S. Zavaleta & E. M. Wolkovich. 2012 Phenological tracking enables positive species responses to climate change. Ecology 93(8): 1765-1771. (article // data)
Pau, S, Gillespie, T.W., & E.M. Wolkovich. 2012. Dissecting NDVI-species richness relationships in Hawaiian dry forests. Journal of Biogeography 39(9): 1678-1686. (pdf // supp)
Davies, T.J., Kraft, N.B.J., Salamin, N. & E. M. Wolkovich. 2012. Incompletely resolved phylogenetic trees inflate estimates of phylogenetic conservatism. Ecology 93(2): 242-247. (abstract)
Wainwright, C.E., Wolkovich, E.M. & E.E. Cleland. 2012. Seasonal priority effects: implications for invasion and restoration in a semi-arid system. Journal of Applied Ecology, 49(1): 234-241. (pdf, recommended by Faculty of 1000)
Craine, J. M., E. M. Wolkovich, & E. G. Towne. 2012. The roles of shifting and filtering in generating community-level flowering phenology. Ecography 35. (pdf)
Cook, B. I., Wolkovich, E. M. & 17 co-authors. 2012. Ecosystems. Sensitivity of spring phenology to warming across temporal and spatial climate gradients in two independent databases. (pdf // PEP & NECTAR data)


Craine, J. M., Wolkovich, E. M., Towne, E. G. & S. W. Kembel. 2011. Flowering phenology as a functional trait. New Phytologist 193: 673-682. (pdf)
Wolkovich, E.M. & E.E. Cleland. 2011. The phenology of plant invasions: A community ecology perspective. Frontiers in Ecology & the Environment 9(5): 287-294. (pdf // data from Project Budburst, featured on Project Biodiversify)
Pau*, S., Wolkovich*, E.M., Cook, B.I., Davies, T.J., Kraft, N.J.B., Bolmgren, K., Betancourt, J. & E.E. Cleland. 2011. Predicting phenology by integrating ecology, evolution and climate science. *Both authors contributed equally to this work. Global Change Biology 17: 3633-3643. (pdf // data)
Wilson, E.E. & E. M. Wolkovich. 2011. Scavenging: How carnivores and carrion structure communities. Trends in Ecology and Evolution 26(3): 129-135. *Both authors contributed equally to this work. (pdf // data & code)
Firn, J. & 32 co-authors (Wolkovich, E.M. co-author) 2011. Abundance of introduced species at home predicts abundance away in herbaceous communities. Ecology Letters 14(3):247-281. (Publication from Nutrient Network. Assisted with writing, editing and statistical analyses.) (pdf)
Nicholas, K. A., Wolkovich, E. M. & L. A. Webb. 2011. Trends in climate and phenological changes in California and Australia. American Journal of Enology and Viticulture 62(3): 395A-395A. (pdf)


Wolkovich E.M. 2010. Non-native grass litter enhances grazing arthropod assemblages by increasing native shrub growth. Ecology 91(3): 756-766. (pdf // data)
Wolkovich E.M., Lipson, D.A., Virginia, R.A., Bolger, D.T., & K.L. Cottingham. 2010. Grass invasion causes rapid increases in ecosystem carbon and nitrogen storage in a semi-arid shrubland. Global Change Biology 16(4): 1352-1365. (pdf // data)
Wolkovich, E.M. 2010 Defining and re-defining invasion biology (book review). Journal of Vegetation Science 21(4): 804-806. (pdf)


Wolkovich E.M., D.T. Bolger & D.A. Holway. 2009. Complex responses to invasive grass litter by ground arthropods in a Mediterranean shrub ecosystem. Oecologia 161(4): 697-708. (pdf // data)
Wolkovich, E. M., D.T. Bolger & K.L. Cottingham. 2009. Invasive grass litter facilitates native shrubs through abiotic effects. Journal of Vegetation Science 20(6): 1121-1132. (pdf // data)


Buchholtz, E.A., Wolkovich E.M. & Cleary, R.J. 2005. Vertebral osteology and complexity in Lagenorhynchus acutus (Delphinidae) with comparison to other Delphinoid genera. Marine Mammal Science 21:411-428. (link)

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